An Evolutionary Question solution
by Kegler

The dropped letters (shown below in red) spell PORK SAUSAGE, the MISSING LINK from 18 Across.

An Evolutionary Question solution grid

Across
1IDLES (IS LED anag)
5P(ARLO)R
9SWANg (rev)
11CU(PC + A)KE
12AN + T
13C + RAM
14A + L(ARM + CL)OCK
16C(L)OD
17WHIR (1st letters)
18MISSING LINK
21A + NTE (NET anag)
25P + ASS
27aGREE + N(L)IGHT
29THUS (SHUT anag)
30NOD (rev)
31R(AT + H)OLE
32KEy + PT
33A(WH)IR + L
34RI(N)SE
Down
1ISAAC (hidden rev)
2DO(WELL)ING
3LA + RV + A
4SCAMP + I
5PA(UN)CH (NU rev; CHAP anag)
6AP[T/p]LY
7L + ARCH
8R (EM) + ARK (ME rev)
10NE(R)D (DEN rev)
13S(C) + OWL
15K + INGS + HIPS (SING anag)
18MANT(R)A + P
19S + EEKS
20GUI(L)DE + R
22T + RUTH (HURT anag)
23TENSOR (STONER anag)
24K + NO + L + L
25PIN + K (NIP rev)
26ESTATE (hidden)
28GLEAN (ANGLE anag)